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Family

Cycloteuthidae

Naef, 1923

Cycloteuthids are mesopelagic squids of moderate size, the largest reaches about 60 cm ML, with a cosmopolitan distribution in tropical and subtropical waters (Nesis, 1982). In Cycloteuthis the mantle and fins are drawn-out into a slender tail. A tail is lacking in Discoteuthis and the large, oval (disc-like) combined fins extend virtually the full length of the mantle. Very little is known about the biology of any species in the family.

An oegopsid ...

  • with triangular funnel locking-apparatus.
  • with mantle component of funnel locking-apparatus not reaching anterior margin of mantle.

An oegopsid ...

  • with triangular funnel locking-apparatus.
  • with mantle component of funnel locking-apparatus not reaching anterior margin of mantle.

This family is represented by seven species within two genera

  • Cycloteuthis
    • Cycloteuthis akimushkini (Filippova, 1968:119)
    • Cycloteuthis sirventi (Joubin, 1919:1)
  • Discoteuthis
    • Discoteuthis discus (Young and Roper, 1969:5)
    • Discoteuthis lacinosa ((Young and Roper, 1969:9)

Nominal Genus-Level Taxa

Cycloteuthis Joubin, 1919:1.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Young and Roper (1969:2)
TYPE SPECIES. -- Cycloteuthis sirventi Joubin, 1919 by monotypy

Discoteuthis Young and Roper, 1969:5.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Nesis (1987:240)]
TYPE SPECIES. -- Discoteuthis discus Young and Roper, 1969 by original designation

Nominal Species-Level Taxa (As Introduced Binomial)

Cycloteuthis akimushkini Filippova, 1968:119, figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:240)]
TYPE REPOSITORY. -- ZMMGU Holotype
TYPE LOCALITY. -- Indian Ocean (9∞57'S, 91∞31'E)

Discoteuthis discus Young and Roper, 1969:5, pls 3-4, 7-9.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:240)]
TYPE REPOSITORY. -- NMNH Holotype 576214 [fide Roper and Sweeney (1978:6)]
TYPE LOCALITY. -- 03∞50'N, 02∞37'W (Atlantic Ocean)

Discoteuthis laciniosa Young and Roper, 1969:9, pls 5-9.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:241)]
TYPE REPOSITORY. -- NMNH Holotype 576215 [fide Roper and Sweeney (1978:8)]
TYPE LOCALITY. -- 33∞04'N, 64∞37'W (Atlantic Ocean)

Cycloteuthis sirventi Joubin, 1919:1, figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Young and Roper (1969:2)
TYPE REPOSITORY. -- MOM Holotype [station 3213] [fide Belloc (150:5)]
TYPE LOCALITY. -- 30∞45'40"N, 25∞47'W, off Madeira

The features that unites these two genera is the similar triangular shape of the funnel locking-apparatus and the position of the mantle component. Other than these unusual features, there is little morphological evidence for their close relationship. Molecular evidence (Carlini, 1998; Lindgren, 2010) supports their close relationship. A more recent study, however, found the two genera only distantly related (Lindgren, 2012).

  1. Arms
    1. Arm suckers biserial.
    2. Buccal-crown connectives attach to ventral borders of Arms IV.
  2. Tentacles
    1. Tentacular club with quadraserial suckers.
  3. Funnel/mantle locking apparatus
    1. Funnel component with sub-triangular groove.
    2. Mantle component does not reach anterior mantle margin.

Comments

The genera are easily separated by body and fin shape (see title illustrations). In addition the presence of a large visceral photophore only in Cycloteuthis allows genera to be separated when badly damaged.

  • Carlini, D. B. 1998. The phylogeny of coleoid cephalopods inferred from molecular evolutionary analyses of the cytochrome oxidase I, muscle actin, and cytoplasmic actin genes. Ph.D. diss. Coll. William and Mary, 273 pp.
  • Lindgren, A.R. 2010.Molecular inference of phylogenetic relationships among Decapodiformes (Mollusca: Cephalopoda) with special focus on the squid Order Oegopsida. Mol. Phylogenet. Evol. 56: 77-90.
  • Lindgren, A. R., M. S. Pankey, F. G. Hochberg and T. H. Oakley. 2012. A multi-gene phylogeny of Cephalopoda supports convergent morphological evolution in association with multiple habitat shifts in the marine environment. BMC Evolutionary Biology 2012, 12:129-144.
  • Nesis, K. N. 1982. Abridged key to the cephalopod mollusks of the world's ocean. 385+ii pp. Light and Food Industry Publishing House, Moscow. (In Russian.). Translated into English by B. S. Levitov, ed. by L. A. Burgess (1987), Cephalopods of the world. T. F. H. Publications, Neptune City, NJ, 351pp.
  • Young, R. E. and C. F. E. Roper. 1969. A monograph of the Cephalopoda of the North Atlantic: The family Cycloteuthidae. Smithsonian Contributions to Zoology No. 5:1-24.
  • Young, Richard E. 2019. Cycloteuthidae Naef, 1923 . Version 26 March 2019 (under construction). http://tolweb.org/Cycloteuthidae/19412/2019.03.26 in The Tree of Life Web Project, http://tolweb.org/

Taxonomy

Family
Cycloteuthidae
Naef, 1923
↔ Sibling Groups (2)